The Opisthothelae are spiders within the order Araneae, consisting of the Mygalomorphae and the Araneomorphae, but excluding the Mesothelae. The Opisthothelae are sometimes presented as an unranked clade[2][3] and sometimes as a suborder of the Araneae.[2] In the latter case, the Mygalomorphae and Araneomorphae are treated as infraorders.

Temporal range: Triassic to present
Missulena bradleyi
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Subphylum: Chelicerata
Class: Arachnida
Order: Araneae
Suborder: Opisthothelae
Pocock, 1892[1]

The fairly recent creation of this taxon has been justified by the requirement to distinguish these spiders from the Mesothelae, which display many more primitive characteristics. Those that distinguish between the Mesothelae and Opisthothelae are:

  • The tergite plates on the abdomen of Mesothelae but absent in Opisthothelae
  • The almost total absence of ganglia in the abdomen of Opisthothelae
  • The almost median position of the spinnerets in the Mesothelae compared with the hindmost position of those of the Opistothelae

Among the Opisthothelae, the fangs of the Mygalomorphae point straight down in front of the mouth aperture and only allow the spider to grasp its prey from above and below, whereas in the Araneomorphae, they face one another like pincers, allowing a firmer grip. Lampshade spiders (family Hypochilidae) show some characteristics of Araneomorphae despite being mygalomorphs and have fangs that can move diagonally.[4] Distinguishing araneomorphs and mygalomorphs on first inspection is difficult unless the specimens are large enough to permit immediate examination of the fangs.


  1. Dunlop, Jason A. & Penney, David (2011). "Order Araneae Clerck, 1757" (PDF). In Zhang, Z.-Q. (ed.). Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa. Auckland, New Zealand: Magnolia Press. ISBN 978-1-86977-850-7. Retrieved 2015-10-31.
  2. Scientific name: Opisthothelae in Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. http://taxonomicon.taxonomy.nl/. Access date: 8 December 2010
  3. Coddington, J.A. (2005). "Phylogeny and Classification of Spiders". In Ubick, D.; Paquin, P.; Cushing, P.E.; Roth, V. (eds.). Spiders of North America: an identification manual (PDF). American Arachnological Society. pp. 18–24. ISBN 0-9771439-0-2. Retrieved 2008-10-12.
  4. Brunetta, Leslie; Craig, Catherine L. (2010). Spider Silk. New Haven and London: Yale University Press.

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